Trilobites of the Wheeler Shale Formation





Altiocculus harrisi

Alokistocare sp.

Asaphiscus wheeleri

Baltagnostus eurypyx

Bathyocos housensis

Bathyuriscus fimbriatus

Bolaspidella housensis

Bolaspidella wellsvillensis

Brachyaspidion microps

Brachyaspidion sulcatum
Elrathia kingii (or kingi)

Elrathina wheeleri = Ptychoparella wheeleri

Hypagnostus parvifrons

Jenkinsonia varga

Modocia brevispina

Modocia laevinucha

Modocia typicalis

Olenoides expansus

Olenoides nevadensis
Onymagnostus seminula

Peronopsis amplaxis

Peronopsis bidens

Peronopsis fallax

Peronopsis gaspensis

Peronopsis intermedius

Peronopsis interstrictus = Itagnostus interstrictus

Peronopsis montis

Peronopsis segmenta

Ptychagnostus atavus

Ptychagnostus gibbus

Ptychagnostus germanus

Ptychagnostus occultatus

Ptychagnostus seminula

Ptychoparella wheeleri

Zacanthoides sp.




Altiocculus sp

Asaphiscus wheeleri

Bathyuriscus sp.

Bolaspidella drumensis

Bolaspidella sp

Elrathia kingii

Elrathia sp

Glyphaspis concavus

Modecia sp

Olenoides expansus

Olenoides serratus

Peronopsis amplaxis

Peronopsis fallax

Peronopsis gaspensis

Peronopsis interstricta

Peronopsis montis

Peronopsis segmenta

Ptychagnostus atavus

Ptychagnostus gibbus

Ptychagnostus intermedius

Ptychagnostus seminula

Ptychoparella sp.

Spencella sp.

Tonkinella breviceps

Zacanthoides divergens

Zacanthoides sp.

Burgess Shale type fossils found with Trilobites of the Wheeler Formation:

Choia sp sponge
Gogia spiralis Eocrinoid
Margaretia dorus sponge
Branchiocaris Phyllocarid


The Drum Mountains portion of the Wheeler are shallow facies and is equivalent to the Pierson Cove Formation. The Elrathia in the Drums are for the most part Elrathia marjumi, although some appear to be an intermediate species between Elrathia marjumi and Elrathia kingii. There is much debate on where the boundary between the Pierson Cove and the Wheeler Formation actually is.





Allison, P.A.; Brett, C.E.; Liddell, W.D.; Wright, E.; 1995.

Repetitive tapho, bio- and litho facies, within the Middle Cambrian Swazey

Limestone-Wheeler Shale transition of Utah. Geological Society of America,

Abstracts with Programs 27(6):374


Babcock, L. E., Rees, M. N., Robison, R. A., Langenburg, E. S., and Peng, S. (2004) Potential Global Standard Stratotype-section and Point (GSSP) for a Cambrian stage boundary defined by the first appearance of the trilobite Ptychagnostus atavus, Drum Mountains, Utah, USA. Geobios 37, pp. 149-158


Gaines, Robert R.; 2000.

Ichnofabric, taphonomy and soft-bodied preservation in the Middle Cambrian

Wheeler Shale, Millard County, Utah. PaleoBios 20(Suppl.1):1-3 le


Gaines, Robert R.; Droser, Mary L.; Kennedy, Martin J.; 2001.

A diagenetic trigger of exceptional preservation in the Wheeler Shale

(Middle Cambrian, Utah). PaleoBios 21(Suppl.1):1-4 S


Gaines, Robert R.; Droser, Mary L.; Kennedy, Martin J.; 2001.

Taphonomy of soft-bodied preservation and ptychopariid Lagerstätte in the

Wheeler Shale (Middle Cambrian), House Range, USA; controls and

implications. PaleoBios, 21(Suppl.2):1:55


Rigby, J. Keith; Gunther, Lloyd F.; Gunther, Val G.; 1983.

Quarries for Middle Cambrian fossils in the Wheeler and Marjum formations

of western Utah. Geological Society of America, Abstracts with Programs



Robison, Richard Ashby; 1962.

Late Middle Cambrian Faunas from the Wheeler and Marjum Formations of

Western Utah. PhD Thesis, University of Texas at Austin


Robison, Richard Ashby; 1971.

Additional Middle Cambrian Trilobites From the Wheeler Shale of Utah.

Journal of Paleontology 45(5):796-804


Rogers, J.C.; 1984.

Depositional environments and paleoecology of two quarry sites in the

Middle Cambrian Marjum and Wheeler Formations, House Range, Utah. Brigham

Young University Geology studies 31, 97-115.


Schneider, Loren P.; 2000.

The sequence stratigraphy of the Middle Cambrian Wheeler Formation in the

Drum Mountains of west central Utah. Monograph.


Sundberg, Frederick A.; 1994.

Corynexochida and Ptychopariida (Trilobita,Arthropoda) of the Ehmaniella

Biozone (Middle Cambrian), Utah and Nevada. Natural History Museum of

Los Angeles County - Contributions in Science, No.446


White, William W. III; 1973.

Depositional environments of Wheeler Formation, Drum Mountains, Millard

County, Utah. The American Association of Petroleum Geologists Bulletin 57(5)

Also see: Fossils of the House Range in Utah


*researched and compiled by James Cook, Andrew Milner and Kevin D. Brett

Western Trilobites Association Home:
Trilobite Genera / Species I Trilobite Formations I Trilobite Locations