Burgess Shale fauna:

 

LIST OF ALL SPECIES (in alphabetical order):

 

Acrothyra gregaria (Walcott, 1924a)  (Brachiopoda)

Actaeus armatus (Simonetta, 1970)  (Arthropoda)

Alalcomenaeus cambricus (Simonetta, 1970)  (Arthropoda)

Aluta?  sp. indet.  (Arthropoda)

Amiella ornata (Walcott, 1911d(Enigmata)

Amiskwia sagittiformis (Walcott, 1911d)  (Enigmata)

Ancalagon minor (Walcott, 1911d)  (Priapulida)

Anomalocaris canadensis (Whiteaves, 1892) 1 (Enigmata)

Anomalocaris nathorsti (Walcott, 1911a) (Enigmata)

Aysheaia pedunculata (Walcott, 1911d)  (Onychophora)

Banffia constricta (Walcott, 1911d) (?Polychaeta)

Bidentia difficilis Walcott, 1912a = Leanchoilia superlata Walcott, 1912a (Arthropoda)

Bosworthia gyges Walcott, 1919  (Rhodophyta)

Bosworthia simulans Walcott, 1919  (Rhodophyta)

Branchiocaris pretiosa (Resser, 1929)  (Arthropoda)

Burgessia bella Walcott, 1912a (Arthropoda)

Burgessochaeta setigera Walcott, 1911d  (Polychaeta)

Cambrorhytium fragilis (Walcott, 1911d)  (?Cnidaria)

Cambrorhytium major (Walcott, 1908a)  (?Cnidaria)

Canadaspis obesa Simonetta and Delle Cave, 1975= Canadaspis perfecta  (Walcott, 1912a)  (Arthropoda)

Canadaspis ovalis (Walcott, 1912a)  (Arthropoda)

Canadaspis perfecta  (Walcott, 1912a)  (Arthropoda)

Canadia grandis Walcott, 1931 = Canadia spinosa Walcott, 1911d (Polychaeta)

Canadia regularis Walcott, 1911d = Canadia spinosa Walcott, 1911d (Polychaeta)

Canadia spinosa Walcott, 1911d  (Polychaeta)

Canistrumella alternata Rigby, 1986  (Porifera)

Capsospongia undulata (Walcott, 1920)  (Porifera)

Carnarvonia venosa Walcott, 1912a  (Arthropoda)

Chancelloria eros Walcott, 1920 (Enigmata)

Chancia palliseri (Walcott, 1908b)  (Arthropoda)

Chaunograptus scandens Ruedemann, 1931 (Hemichordata)

Choia carteri Walcott, 1920 (Porifera)

Choia ridleyi Walcott, 1920  (Porifera)

Crumillospongia biporosa Rigby, 1986  (Porifera)

Crumillospongia frondosa Walcott, 1920 (Porifera)

Dalyia nitans Walcott, 1919  (Rhodophyta)

Dalyia racemata Walcott, 1919  (Rhodophyta)
Darwin's Dilemma Blog, Morris Conway Bibliography, retrieved July 14, 2013.

Diagoniella hindei Walcott, 1920  (Porifera)

Dictyophycus gracilis Ruedemann, 1931  (Possible Algae)

Dinomischus isolatus Conway Morris, 1977b (Enigmata)

Diraphora bellicostata (Walcott, 1924a)  (Brachiopoda)

Echmatocrinus brachiatus Sprinkle, 1973  (Echinodermata)

Ehmaniella burgessensis Rasetti, 1951  (Arthropoda)

Ehmaniella waptaensis Rasetti, 1951  (Arthropoda)

Eiffelia globosa Walcott, 1920  (Porifera)

Eldonia ludwigi Walcott, 1911c  (Echinodermata)

Elrathia permulta (Walcott, 1918b)  (Arthropoda)

cf. Elrathina brevifrons Rasetti, 1951  (Arthropoda)

Elrathina cordillerae (Rominger, 1887)  (Arthropoda)

Emeraldella brocki Walcott, 1912a (Arthropoda)

Emeraldella micrura Walcott, 1912a = Leanchoilia superlata Walcott, 1912a (Arthropoda)

Emeraldoides problematicus Simonetta, 1964) = Emeraldella brocki Walcott, 1912a (Arthropoda)

Eurysaces pielus Simonetta and Delle Cave, 1975) = Odaria alata Walcott, 1912a  (Arthropoda)

Falospongia alata Rigby, 1986  (Porifera)

Fasciculus vesanus Simonetta and Delle Cave, 1978a (Ctenophora)

Fieldia lanceolata Walcott, 1912a  (Priapulida)

Fieldospongia bellilineata (Walcott, 1920) (Porifera)?Gogia radiata Sprinkle, 1973  (Echinodermata)

Habelia? brevicaudata Simonetta, 1964  (Arthropoda)

Habelia optata Walcott, 1912a  (Arthropoda)

Habelia expansa Walcott, 1918  (Arthropoda)

Halichondrites elissa Walcott, 1920 (Porifera)

Hallucigenia sparsa (Walcott, 1911d)  (Onychophora)

Hamptonia bowerbanki Walcott, 1920  (Porifera)

Hanburia gloriosa Walcott, 1916b  (Arthropoda)

Haplophrentis carinatus (Matthew, 1899) (Hyolitha)

Hazellia conferta Walcott, 1920 (Porifera)

Hazellia crateria Rigby, 1986 (Porifera)

Hazellia delicatula Walcott, 1920  (Porifera)

Hazellia dignata (Walcott, 1920)  (Porifera)

Hazellia grandis Walcott, 1920  (Porifera)

Hazellia luteria Rigby, 1986  (Porifera)

Hazellia nodulifera Walcott, 1920  (Porifera)

Hazellia obscura Walcott, 1920  (Porifera)

Hazellia palmata Walcott, 1920  (Porifera)

Helmetia expansa Walcott, 1918a  (Arthropoda)

Houghtonites gracilis (Walcott, 1912a)  (Arthropoda)

Hurdia dentata Simonetta and Delle Cave, 1975  (Enigmata)

Hurdia triangulata Walcott, 1912a  (Enigmata)

Hurdia victoria Walcott, 1912a  (Enigmata)

Hymenocaris? circularis Walcott, 1912a) = Burgessia bella Walcott, 1912a (Arthropoda)

Hymenocaris obliqua Walcott, 1912a = Canadaspis perfecta  (Walcott, 1912a)  (Arthropoda)

?Hyolithes carinatus  = Haplophrentis carinatus (Matthew, 1899) (Hyolitha)

Insolicorypha psygma Conway Morris 1979a  (Polychaeta)

Isoxys acutangulus (Walcott, 1908a) (Arthropoda)

Isoxys longissimus Simonetta and Delle Cave, 1975 (Arthropoda)

Kootenia burgessensis Resser, 1942  (Arthropoda)

Laggania cambria Walcott, 1911a = Anomalocaris nathorsti (Walcott, 1911a) (Enigmata)

Leanchoilia amphiction Simonetta, 1970 = Leanchoilia superlata Walcott, 1912a (Arthropoda)

Leanchoilia major Walcott, 1931 = Leanchoilia superlata Walcott, 1912a (Arthropoda)

Leanchoilia persephone Simonetta, 1970 = Leanchoilia superlata Walcott, 1912a (Arthropoda)

Leanchoilia protogonia Simonetta, 1970) = Leanchoilia superlata Walcott, 1912a (Arthropoda)

Leanchoilia superlata Walcott, 1912a (Arthropoda)

Lecythioscopa simplex Walcott, 1931  (?Priapulida)

Leptomitus lineatus (Walcott, 1920)  (Porifera)

Lingulella waptaensis Walcott, 1924a  (Brachiopoda)

Louisella pedunculata Walcott, 1911d  (Priapulida)

Mackenzia costalis Walcott, 1911c  (Cnidaria)

Margaretia dorus Walcott, 1931 (Chlorophyta)

Marpolia spissa Walcott, 1919  (Cyanobacteria)

Marrella splendens Walcott, 1912a  (Arthropoda)

Metaspriggina walcotti Simonetta and Insom, 1993 (Chordata)

Micromitra burgessensis Resser, 1938  (Brachiopoda)

Miskoia placida Walcott, 1931 = Ottoia prolifica Walcott, 1911d (Priapulida)

Miskoia preciosa Walcott, 1911d = Louisella pedunculata Walcott, 1911d  (Priapulida)

Molaria spinifera Walcott, 1912a  (Arthropoda)

Moleculopina mammilata (Walcott, 1920)  (Porifera)

Mollisonia rara Walcott, 1912a (Arthropoda)

Mollisonia symmetrica Walcott, 1912a (Arthropoda)

Morania confluens Walcott, 1919 (Cyanobacteria)

Morania  elongata Walcott, 1919 (Cyanobacteria)

Morania  fragmenta Walcott, 1919 (Cyanobacteria)

Morania? frondosa Walcott, 1919 (Cyanobacteria)

Morania? globosa Walcott, 1919 (Cyanobacteria)

Morania  parasitica Walcott, 1919 (Cyanobacteria)

Morania? reticulata Walcott, 1919 (Cyanobacteria)

Naraoia compacta Walcott, 1912a (Arthropoda)

Naraoia  halia Simonetta and Delle Cave, 1975) = Naraoia compacta Walcott, 1912a (Arthropoda)

Naraoia  pammon Simonetta and Delle Cave, 1975 = Naraoia compacta Walcott, 1912a (Arthropoda)

Naraoia spinifer Walcott, 1931  (Arthropoda)

Nathorstia transitans Walcott, 1912a)  = Olenoides serratus (Rominger, 1887) (Arthropoda)

Nectocaris pteryx Conway Morris, 1976b  (Enigmata)

Nisusia burgessensis Walcott, 1924a  (Brachiopoda)

Odaria alata Walcott, 1912a  (Arthropoda)

Odontogriphus omalus Conway Morris, 1976a (?Lophophorata)

Oesia disjuncta Walcott, 1911d  (Enigmata)

Olenoides serratus (Rominger, 1887) (Arthropoda)

Opabinia regalis Walcott, 1912a  (Enigmata)

Oryctocephalus burgessensis Resser, 1938 (Arthropoda)

Oryctocephalus matthewi Rasetti, 1951  (Arthropoda)

Oryctocephalus reynoldsi Reed, 1899  (Arthropoda)

Oryctocephalus sp. indet.  (Arthropoda)

Ottoia prolifica Walcott, 1911d  (Priapulida)

"Ottoia" tenuis Walcott, 1911d  (Hemichordata)

Pagetia bootes Walcott, 1916b  (Arthropoda)

Parahabelia rara Simonetta,1964) = Mollisonia rara Walcott, 1912a (Arthropoda)

Parkaspis decamera Rasetti, 1951  (Arthropoda)

Paterinazenobia (Walcott, 1924a)  (Brachiopoda)

Peronochaeta dubia (Walcott, 1911d)  (Polychaeta)

Peronopsis montis (Matthew, 1899)  (Arthropoda)

Perspicaris dicynna (Simonetta and Delle Cave, 1975) (Arthropoda)

Perspicaris recondita Briggs, 1977  (Arthropoda)

Peytoia nathorsti Walcott, 1911a) = Anomalocaris nathorsti (Walcott, 1911a) (Enigmata)

Pikaia gracilens Walcott, 1911d  (Chordata)

Pirania muricata Walcott, 1920  (Porifera)

"Platydendron ovale"  Simonetta and Delle Cave, 1978a (Enigmata)

Plenocaris plena (Walcott, 1912a)  (Arthropoda)

Pollingeria grandis Walcott, 1911d  (Enigmata)

Portalia mira Walcott, 1918a  (Enigmata)

Priscansermarinus barnetti Collins and Rudkin, 1981  (Arthropoda)

Proboscicaris agnosta Rolfe, 1962  (Enigmata)

Proboscicaris ingens, Rolfe, 1962 (Enigmata)

Proboscicaris obtusa Simonetta and Delle Cave, 1975  (Enigmata)

Protospongia hicksi Hinde, 1887  (Porifera)

Ptychagnostus praecurrens (Westergaard, 1936) (Arthropoda)

Sanctacaris uncata Briggs and Collins, 1988  (Arthropoda)

Sarotrocercus oblita Whittington, 1981b  (Arthropoda)

Scenella amii (Matthew, 1902)  (?Mollusca)

Scolecofurca rara Conway Morris, 1977d  (?Priapulida)

Selkirkia columbia Conway Morris, 1977d  (Priapulida)

Selkirkia major Walcott, 1911d = Selkirkia columbia Conway Morris, 1977d  (Priapulida)

Sentinelia draco Walcott, 1920  (Porifera)

Sidneyia inexpectans Walcott, 1911a  (Arthropoda)

Skania fragilis Walcott, 1931  (Arthropoda)

Spencella sp. indet. 1 (Rasetti, 1951)  (Arthropoda)

Spencella sp. indet. 2 (Rasetti, 1951)  (Arthropoda)

Sphaerocodium? cambria Walcott, 1919  (Rhodophyta)

Sphaerocodium? praecursor Walcott, 1919  (Rhodophyta)

Stephanospongia magnipora Rigby, 1986  (Porifera)

Stephenocolex argutus Conway Morris, 1979a  (Polychaeta)

Takakkawia lineata Walcott, 1920 (Porifera)

Tegopelte gigas Simonetta and Delle Cave, 1975  (Arthropoda)

Thaumaptilon walcotti Conway Morris, 1993  (Cnidaria)

Thelxiope palaeothallasia Simonetta and Delle Cave, 1975 (Arthropoda)

Triplagnostus burgessensis Rasetti, 1951) = Ptychagnostus praecurrens (Westergaard, 1936) (Arthropoda)

Tubulella  (?Polychaeta)

Tuzoia burgessensis Resser, 1929  (Arthropoda)

Tuzoia canadensis Resser, 1929  (Arthropoda)

Tuzoia? parva (Walcott, 1912a)  (Arthropoda)

Tuzoia praemorsa Resser, 1929  (Arthropoda)

Tuzoia retifera Walcott, 1912a  (Arthropoda)

Vauxia bellula Walcott, 1920  (Porifera)

Vauxia densa Walcott, 1920  (Porifera)

Vauxia gracilenta Walcott, 1920  (Porifera)

Vauxia venata Walcott, 1920  (Porifera)

Wahpia mimica Walcott, 1919  (Rhodophyta)

Wahpia virgata Walcott, 1919  (Rhodophyta)

Walcottidiscus magister Bassler, 1936  (Echinodermata)

Walcottidiscus typicalis Bassler, 1935  (Echinodermata)

Wapkia grandis Walcott, 1920  (Porifera)

Waptia fieldensis Walcott, 1912a  (Arthropoda)

Waputikia ramosa Walcott, 1919  (Rhodophyta)

Wiwaxia corrugata (Matthew, 1899)  (Polychaeta)

Worthenella cambria Walcott, 1911d  (Enigmata)

Yohoia tenuis Walcott, 1912a  (Arthropoda)

Yuknessia simplex Walcott, 1919 (?Chlorophyta)

 

 

 

BURGESS SHALE FAUNAL LISTING (taxonomic listing):

 

            [ ] = approx. # of genera

            ( ) = approx. # of species

            in bold: undescribed

 

                       

Cyanobacteria-Hormogonales (Filamentous Cyanobacteria) [2]:

            Marpolia (1)

                        spissa Walcott, 1919

            Morania (7)

                        confluens Walcott, 1919

                                elongata Walcott, 1919

                        fragmenta Walcott, 1919

                        M? frondosa Walcott, 1919

                        M? globosa Walcott, 1919

                        parasitica Walcott, 1919

                        M? reticulata Walcott, 1919

Algae [8]:

            Chlorophyta (Green Algae):

                        Margaretia (1)

                                    dorus Walcott, 1931

            ?Chlorophyta:

                        Yuknessia (1)

                                    simplex Walcott, 1919

            Rhodophyta (Red Algae):

                        Bosworthia (2)

                                    gyges Walcott, 1919

                                    simulans Walcott, 1919

                        Dalyia (2)

                                    nitans Walcott, 1919

                                    racemata Walcott, 1919

                        Wahpia (2)

                                    mimica Walcott, 1919

                                    virgata Walcott, 1919

                        Waputikia (1)

                                    ramosa Walcott, 1919

            Rhodophyta (Calcareous Algae):

                        Sphaerocodium ?(2)

                                    S? cambria Walcott, 1919

                                    S? praecursor Walcott, 1919

Possible Algae:

            Dictyophycus (1)

                        gracilis Ruedemann, 1931

Phylum Porifera [20]:

            Class Demospongia:

                        Capsospongia (1)

                                    undulata (Walcott, 1920)

                        Choia (2)

                                    carteri Walcott, 1920

                                    ridleyi Walcott, 1920

                        Crumillospongia (2)

                                    biporosa Rigby, 1986

                                    frondosa Walcott, 1920

                        Falospongia (1)

                                    falata Rigby, 1986

                        Fieldospongia (1)

                                    bellilineata (Walcott, 1920)

                        Halichondrites (1)

                                    elissa Walcott, 1920

                        Hamptonia (1)

                                    bowerbanki Walcott, 1920

                        Hazellia (9)

                                    conferta Walcott, 1920

                                    crateria Rigby, 1986

                                    delicatula Walcott, 1920

                                    dignata (Walcott, 1920)

                                    grandis Walcott, 1920

                                    luteria Rigby, 1986

                                    nodulifera Walcott, 1920

                                    obscura Walcott, 1920

                                    palmata Walcott, 1920

                        Leptomitus (1)

                                    lineatus (Walcott, 1920)

                        Moleculopina (1)

                                    mammilata (Walcott, 1920)

                        Pirania (1)

                                    muricata Walcott, 1920

                        Sentinelia (1)

                                    draco Walcott, 1920

                        Takakkawia (1)

                                    lineata Walcott, 1920

                        Vauxia (4)

                                    bellula Walcott, 1920

                                    densa Walcott, 1920

                                    gracilenta Walcott, 1920

                                    venata Walcott, 1920

                        Wapkia (1)

                                    grandis Walcott, 1920

            Class: Hexactinellida (Glass sponges):

                        Diagoniella (1)

                                    hindei Walcott, 1920

                        Protospongia (1)

                                    hicksi Hinde, 1887

                        Stephanospongia (1)

                                    magnipora Rigby, 1986

            Class Calcarea:

                        Canistrumella (1)

                                    alternata Rigby, 1986

                        Eiffelia (1)

                                    globosa Walcott, 1920

Coelenterata [4]:

Phylum Cnidaria:

            ?Class Anthozoa:

                        ?Order Pennatulacea (Sea Pens and Sea Pansies):

                                    Thaumaptilon (1)

                                                walcotti Conway Morris, 1993

                        ?Order Actiniaria (Sea anemones):

                                    Mackenzia  (1)

                                                costalis Walcott, 1911c

            ?Class Hydrozoa

                        ?Suborder Chondrophorina (By-the-wind Sailors)

                                    Gelenoptron (1)

                                                tentaculatum Conway Morris, 1993

?Phylum Cnidaria

            Cambrorhytium (2)

                        fragilis (Walcott, 1911d)

                        major (Walcott, 1908a)

Phylum Ctenophora:

            Fasciculus  (1-maybe 2)

                        vesanus Simonetta and Delle Cave, 1978a

?Superphylum Lophophorata:

            Odontogriphus (1)

                        omalus Conway Morris, 1976a

Superphylum Lophophorata:

            Phylum Brachiopoda [6]:

                        Class Inarticulata:

                                    Acrothyra (1)

                                                gregaria Walcott, 1924a

                                    Lingulella (1)

                                                waptaensis Walcott, 1924a

                                    Micromitra (1)

                                                burgessensis Resser, 1938

                                    Paterina (1)

                                                zenobia (Walcott, 1924a)

                        Class Articulata:

                                    Diraphora (1)

                                                bellicostata (Walcott, 1924a)

                                    Nisusia (1)

                                                 burgessensis Walcott, 1924a

?Phylum Mollusca [3]:

            Class Helcionelloida (Monoplacophora):

                        Scenella (1)

                                    amii (Matthew, 1902)

Phylum Hyolitha (Hyoliths):

            Haplophrentis  (1) (?Hyolithes carinatus)

                        carinatus (Matthew, 1899)

Priapulida [7]:

Phylum Priapulida:

            Ancalagon (priapulid or Acanthocephala) (1)

                        minor (Walcott, 1911d)

            Fieldia (1)

                        lanceolata Walcott, 1912a

            Louisella (1)

                        pedunculata Walcott, 1911d

                                    (=Miskoia preciosa Walcott, 1911d)

            Ottoia (1)

                        prolifica Walcott, 1911d

                                    (=Miskoia placida Walcott, 1931)

            Selkirkia (1)

                        columbia Conway Morris, 1977d

                                    (=S. major Walcott, 1911d)

            Probable priapulids:

                        Lecythioscopa (1)

                                    simplex Walcott, 1931

                        Scolecofurca (1)

                                    rara Conway Morris, 1977d

Phylum Annelida [25]:

            Class Polychaeta:

                        Burgessochaeta (1)

                                    setigera Walcott, 1911d

                        Canadia (1)

                                    spinosa Walcott, 1911d

                                                (=C. regularis Walcott, 1911d and C. grandis Walcott, 1931)

                        Insolicorypha (1)

                                    psygma Conway Morris 1979a

                        Peronochaeta  (1)

                                    dubia (Walcott, 1911d)

                        Stephenocolex  (1)

                                    argutus Conway Morris, 1979a

                        Wiwaxia  (1) (Briggs et al., 1994 classify as a sclerotome-bearing animal)

                                    corrugata (Matthew, 1899)

Other annelids:

                        Banffia (1)

                                    constricta Walcott, 1911d (Briggs et al., 1994 list as an animal not assigned to a major group)

                        Tubulella (not in Briggs et al.,1994)

 Phylum Onychophora [2]:

            Aysheaia (1)

                        pedunculata Walcott, 1911d

            Hallucigenia (1) (? onychophoran)

                        sparsa (Walcott, 1911d)

Phylum Arthropoda [44], including 14 trilobites:

            Primitive:

                        Branchiocaris (1)

                                    pretiosa (Resser, 1929)

                        Marrella (1 - maybe 2, Devereux pers. comm.)

                                    splendens Walcott, 1912a

            Subphylum Crustacea and Crustaceanomorphs:

                        Canadaspis (2)

                                    ovalis (Walcott, 1912a)

                                    perfecta  (Walcott, 1912a) (=Hymenocaris obliqua Walcott, 1912a and Canadaspis obesa Simonetta and Delle Cave, 1975)

                        Carnarvonia (1)

                                    venosa Walcott, 1912a

                        Isoxys (2)

                                    acutangulus (Walcott, 1908a)

                                    longissimus Simonetta and Delle Cave, 1975

                        Odaria  (1)

                                    alata Walcott, 1912a

                                                (=Eurysaces pielus Simonetta and Delle Cave, 1975)

                        Perspicaris (2)

                                    dicynna (Simonetta and Delle Cave, 1975)

                                    recondita Briggs, 1977

                        Plenocaris (1)

                                    plena (Walcott, 1912a)

                        Tuzoia (5)

                                    burgessensis Resser, 1929

                                    canadensis Resser, 1929

                                    T? parva (Walcott, 1912a)

                                    praemorsa Resser, 1929

                                    retifera Walcott, 1912a

                        Waptia (1)

                                    fieldensis Walcott, 1912a

                        Class Ostracoda:

                                    Aluta? (1)

                                                sp. indet. 1

                        Class Cirripedia:

                                    Priscansermarinus (1) (?Lepidomorph barnacle)

                                                barnetti Collins and Rudkin, 1981

            Subphylum Arachnomorpha:

                        Class Trilobita:

                                    Chancia (1)

                                                palliseri (Walcott, 1908b)

                                    Ehmaniella (2)

                                                burgessensis Rasetti, 1951

                                                waptaensis Rasetti, 1951

                                    Elrathia (1)

                                                permulta (Walcott, 1918b)

                                    Elrathina (2)

                                                cf. E. brevifrons Rasetti, 1951

                                                cordillerae (Rominger, 1887)

                                    Hanburia (1)

                                                gloriosa Walcott, 1916b

                                    Kootenia (1)

                                                burgessensis Resser, 1942

                                    Naraoia (2)

                                                compacta Walcott, 1912a (=N. pammon Simonetta and Delle Cave, 1975 and N. halia Simonetta and Delle Cave, 1975)

                                                spinifer Walcott, 1931

                                    Olenoides (1)

                                                serratus (Rominger, 1887) (=Nathorstia transitans Walcott, 1912a)

                                    Oryctocephalus (4)

                                                burgessensis Resser, 1938

                                                matthewi Rasetti, 1951

                                                reynoldsi Reed, 1899

                                                sp. indet. 1

                                    Pagetia (1)

                                                bootes Walcott, 1916b

                                    Parkaspis (1)

                                                decamera Rasetti, 1951

                                    Peronopsis (1)

                                                montis (Matthew, 1899)

                                    Ptychagnostus (1)

                                                praecurrens (Westergaard, 1936) (=Triplagnostus burgessensis Rasetti, 1951)

                                    Spencella (2)

                                                sp. indet. 1 (Rasetti, 1951)

                                                sp. indet. 2 (Rasetti, 1951)

                                    Tegopelte (1)

                                                gigas Simonetta and Delle Cave, 1975

                        Class Chelicerata:

                                    Sanctacaris (1)

                                                uncata Briggs and Collins, 1988

                        Other Arthropods:

                                    Actaeus (1)

                                                armatus Simonetta, 1970

                                    Alalcomenaeus (1)

                                                cambricus Simonetta, 1970

                                    Burgessia (1)

                                                bella Walcott, 1912a (=Hymenocaris? circularis Walcott, 1912a)

                                    Emeraldella (1)

                                                brocki Walcott, 1912a (=Emeraldoides problematicus Simonetta, 1964)

                                    Habelia (3)

                                                H? brevicaudata Simonetta, 1964

                                                optata Walcott, 1912a

                                                expansa Walcott, 1918

                                    Helmetia (1)

                                                expansa Walcott, 1918a

                                    Houghtonites (1)

                                                gracilis (Walcott, 1912a)

                                    Leanchoilia (1)

                                                superlata Walcott, 1912a (=Bidentia difficilis Walcott, 1912a; Emeraldella micrura Walcott, 1912a; Leanchoilia major Walcott, 1931; L. amphiction Simonetta, 1970; L. persephone Simonetta, 1970; L. protogonia Simonetta, 1970)

                                    Molaria (1)

                                                spinifera Walcott, 1912a

                                    Mollisonia (2)

                                                rara Walcott, 1912a (=Parahabelia rara Simonetta,1964)

                                                symmetrica Walcott, 1912a

                                    Sarotrocercus (1)

                                                oblita Whittington, 1981b

                                    Sidneyia (1)

                                                inexpectans Walcott, 1911a

                                    Skania (1)

                                                fragilis Walcott, 1931

                                    Thelxiope (1)

                                                palaeothallasia Simonetta and Delle Cave, 1975

                                    Yohoia (1)

                                                tenuis Walcott, 1912a 

Phylum Echinodermata [5]:

            Class Cystoidea:

                        ?Gogia (1)

                                    radiata Sprinkle, 1973

            Class Crinoidea:

                        Echmatocrinus (1)

                                    brachiatus Sprinkle, 1973

            Class Edrioasteroidea:

                        Walcottidiscus (2)

                                    magister Bassler, 1936

                                    typicalis Bassler, 1935

            Class Holothuroidea:

                        Eldonia  (1)

                                    ludwigi Walcott, 1911c 

Phylum Chordata:

            Metaspriggina  (1)

                        walcotti Simonetta and Insom, 1993

            Pikaia (1)

                        gracilens Walcott, 1911d

Phylum Hemichordata

            Class Enteropneusta?:

                        "Ottoia"  (1)

                                    tenuis Walcott, 1911d

            Class Graptolithina?

                        Chaunograptus (1)

                                    scandens Ruedemann, 1931

 

Enigmata (animals not assigned to major groups):

            Anomalocarids [3?]:

                        Amiella (1)

                                    ornata Walcott, 1911d

                        Anomalocaris (2)

                                    canadensis Whiteaves, 1892

                                    nathorsti (Walcott, 1911a) (=Laggania cambria Walcott, 1911a and Peytoia nathorsti Walcott,                                               1911a)

                        Appendage "F"

                        Hurdia  (3)

                                    dentata Simonetta and Delle Cave, 1975

                                    triangulata Walcott, 1912a

                                    victoria Walcott, 1912a

                        Peytoia (several spp.)

                        Proboscicaris (3)

                                    agnosta Rolfe, 1962

                                    ingens, Rolfe, 1962

                                    obtusa Simonetta and Delle Cave, 1975

            Others:

                        Amiskwia (1)

                                    sagittiformis Walcott, 1911d

                        Dinomischus (1)

                                    isolatus Conway Morris, 1977b

                        Nectocaris (1)

                                    pteryx Conway Morris, 1976b

                        Oesia (1)

                                    disjuncta Walcott, 1911d

                        Opabinia (1)

                                    regalis Walcott, 1912a

                        "Platydendron"  (1)

                                    "Platydendron ovale"  Simonetta and Delle Cave, 1978a

                        Pollingeria  (1)

                                    grandis Walcott, 1911d

                        Portalia (1)

                                    mira Walcott, 1918a

                        Worthenella (1)

                                    cambria Walcott, 1911d

            Scleritome-bearing animals:

                        Chancelloria (sponge?) (1)

                                    eros Walcott, 1920

 

 

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Simonetta, A.M. 1963. Observazion: sugli artropodi non trilobiti della Burgess Shale (Cambriano medio). II. Contributo: I Generi Sidneya [sic] ed Amiella Walcott 1911. Monitore Zoologico Italiano 70: 97-108.

 

Simonetta, A.M. 1970. Studies on non-trilobite arthropods of the Burgess Shale (Middle Cambrian). Palaeontographica Italica 66: 35-45.

 

Simonetta, A.M. 1975. The Cambrian non-trilobite arthropods from the Burgess Shale of British Columbia - a study of their comparative morphology, taxinomy [sic] and evolutionary significance. Palaeontographica Italica 69: 1-37.

 

Siveter, D.J., and Williams, M. 1997. Cambrian bradoriid and phosphatocopid arthropods of North America. Special Papers in Palaeontology, Palaeontological Association, London 57: 1-61.

 

Sprinkle, J., and Collins, D.H. 1998. Revision of Echmatocrinus from the Middle Cambrian Burgess Shale of British Columbia. Lethaia 31: 269-282.

 

Srivastava, D.S., and Lawton, J.H. 1998. Why more productive sites have more species: an experimental test of theory using tree-hole communities. The American Naturalist 152(4): 509-529.

 

Staff, G.M., Stanton, R.J. Jr., Powell, E.N., and Cummins, H. 1986. Time-averaging, taphonomy, and their impact on paleocommunity reconstruction: death assemblages in Texas bays. Geological Society of America Bulletin 97: 428-443.

 

Stanton, R.J. (Jr.), and Dodd, J.R. 1976. The application of trophic structure of fossil communities in paleoenvironmental reconstruction. Lethaia 9: 327-342.

 

Stanton, R.J., and Nelson, P.C. 1980. Reconstruction of the trophic web in paleontology: community structure in the Stone City Formation (Middle Eocene, Texas). Journal of Paleontology 54: 118-135.

 

Storch, V., and Alberti, G. 1985. Ultrastructural investigation of the integument of Tubiluchus philippinensis (Priapulida, Tubiluchidae). Zoologica Scripta 14(4): 265-272.

 

Storch, V., Higgins, R.P., and Morse, M.P. 1989. Ultrastructure of the body wall of Meiopriapulus fijiensis (Priapulida). Transactions of the American Microscopical Society 108(4): 319-331.

 

Strong, D.R. 1997. Quick indirect interactions in intertidal food webs. Trends in Ecology and Evolution 12(5): 173-174.

 

Sun, W.-G., and Hou, X.-G. 1987. Early Cambrian medusae from Chengjiang, Yunnan, China. Acta Palaeontologica Sinica 26(3): 268-279.

 

Tamm, S., and Tamm, S.L. 1995. A giant nerve net with multi-effector synapses underlying epithelial adhesive strips in the mouth of Beroe (Ctenophora). Journal of Neurocytology 24(9): 711-723.

 

Tamm, S.L., and Tamm, S. 1991. Reversable epithelial adhesion closes the mouth of Beroe, a carnivorous marine jelly. Biological Bulletin (Woods Hole) 181(3): 463-473.

 

Trott, T.J. 1998. Gustatory responses of Priapulus caudatus Lamarck, 1816 (Priapulida, Priapulidae): feeding behavior and chemoreception by a living fossil. Marine Behavior and Physiology 31(4): 251-257.

 

Van Der Land, J. 1970. Systematics, zoogeography and ecology of the Priapulida. Zoologische Verhandelingen 112: 1-118.

 

Vannier, J., and Chen, J.-Y. 2000. The Early Cambrian colonization of pelagic niches exemplified by Isoxys (Arthropoda). Lethaia 33: 295-311.

 

Verity, P.G., and Smetacek, V. 1996. Organism life cycles, predation, and the structure of marine pelagic ecosystems. Marine Ecology Progress Series 130: 277-293.

 

Walcott, C.D. 1908. Mount Stephen rocks and fossils. Canadian Alpine Journal 1(2): 232-248.

 

Walcott, C.D. 1911a. Cambrian geology and paleontology II: Middle Cambrian Merostomata. Smithsonian Miscellaneous Collections 57(2): 17-40.

 

Walcott, C.D. 1911b. Cambrian geology and paleontology II: Middle Cambrian holothurians and Medusae. Smithsonian Miscellaneous Collections 57(3): 41-68.

 

Walcott, C.D. 1911c. Cambrian geology and paleontology II: Middle Cambrian annelids. Smithsonian Miscellaneous Collections 57(5): 109-144.

 

Walcott, C.D. 1912a. Cambrian geology and paleontology II: Middle Cambrian Branchiopoda, Malacostraca, Trilobita and Merostomata. Smithsonian Miscellaneous Collections 57(6): 145-228.

 

Walcott, C.D. 1912b. Cambrian of the Kicking Horse Valley, British Columbia. Summary Report, Geological Survey Branch, Department of Mines, Canada, Sessional Paper 26: 188-191.

 

Walcott, C.D. 1918. Cambrian geology and paleontology IV: Appendages of trilobites. Smithsonian Miscellaneous Collections 67(4): 115-216.

 

Walcott, C.D. 1919. Cambrian geology and paleontology IV: Middle Cambrian algae. Smithsonian Miscellaneous Collections 67(5): 217-260.

 

Walcott, C.D. 1920. Cambrian geology and paleontology IV: Middle Cambrian Spongiae. Smithsonian Miscellaneous Collections 67(6): 261-364.

 

Walcott, C.D. 1931. Addenda to descriptions of Burgess Shale Fossils (Posthumous, with explanatory notes by Charles E. Resser). Smithsonian Miscellaneous Collections 85(3): 1-46.

 

Whiteaves, J.F. 1892. Description of a new Genus and Species of phyllocarid Crustacea from the Middle Cambrian of Mount Stephen, B.C. Canadian Record of Science 5(4): 205-208.

 

Whittington, H.B. 1971. The Burgess Shale: history of research and preservation of fossils. Proceedings of the North American Paleontological Convention, Chicago 1: 1170-1201.

 

Whittington, H.B. 1975. The enigmatic animal Opabinia regalis, Middle Cambrian, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society of London B 271: 1-43.

 

Whittington, H.B. 1977. The Middle Cambrian trilobite Naraoia, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society of London B 280: 409-443.

 

Whittington, H.B. 1980b. Exoskeleton, moult stage, appendage morphology and habits of the Middle Cambrian trilobite Olenoides serratus. Palaeontology 23(1): 171-204.

 

Whittington, H.B. 1981. Rare arthropods from the Burgess Shale, Middle Cambrian, British Columbia. Philosophical Transactions of the Royal Society of London B 292: 329-357.

 

Whittington, H.B. 1985. The Burgess Shale. Yale University Press, New Haven: 151 pp.

 

Whittington, H.B., and Briggs, D.E.G. 1982. A new conundrum from the Middle Cambrian Burgess Shale. Third North American Paleontological Convention, Proceedings 2: 573-575.

 

Whittington, H.B., and Briggs, D.E.G. 1985. The largest Cambrian animal, Anomalocaris, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society of London B 309: 569-609.

 

Williams, M., Siveter, D.J., and Peel, J.S. 1996. Isoxys (Arthropoda) from the Early Cambrian Sirius Passet Lagerstätte, north Greenland. Journal of Paleontology 70(6): 947-954.

 

Wills, M.A. 1998. Cambrian and Recent disparity: the picture from priapulids. Paleobiology 24(2): 177-199.

 

Wu, R.S.S., and Shin, P.K.S. 1997. Sediment characteristics and colonization of soft-bottom benthos: a field manipulation experiment. Marine Biology 128: 475-487.

Virtual Fossul Museum - Burgess Shale; Also see Chengjiang Maotianshan Shales and Chengjiang Biota

Yochelson, E.L. 1961. The operculum and mode of life of Hyolithes. Journal of Paleontology 35(1): 152-161.

 

 

 

Burgess taxonomy authors - descriptions of new species (refs. from Briggs et al., 1994):

 

Bassler (1935)

Bassler (1936)

Briggs (1977)

Briggs and Collins (1988)

Briggs, D. E. G., Erwin, D. H., and Collier, F. J. (1994)  The Fossils of the Burgess Shale. Smithsonian Institution Press, Washington, 238 pp.

Collins and Rudkin (1981)

Conway Morris, S. (1976a)

Conway Morris, S. (1976b)

Conway Morris, S. (1977b)

Conway Morris, S. (1977d)

Conway Morris, S. (1979a)

Conway Morris, S. (1993)

Hinde (1887)

Matthew (1899)

Matthew (1902)

Rasetti (1951)

Reed (1899)

Resser (1929)

Resser (1938)

Resser (1942)

Rigby (1986)

Rolfe (1962)

Rominger (1887)

Ruedemann (1931)

Simonetta (1964)

Simonetta (1970)

Simonetta and Delle Cave (1975)

Simonetta and Delle Cave (1978a)

Simonetta and Inson (1993)

Sprinkle (1973)

Walcott (1908a)

Walcott (1908b)

Walcott (1911a)

Walcott (1911c)

Walcott (1911d)

Walcott (1912a)

Walcott, 1916b)

Walcott (1918a)

Walcott (1918b)

Walcott (1919)

Walcott (1920)

Walcott (1924a)

Walcott (1931)

Westergaard (1936)

Whiteaves (1892)

Whittington (1981b)

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